Browse Human Kinome [ Human kinases ]

The kinase complement of the human genome, the kinome, provided a start point for comprehensive investigation of protein phosphorylation. Manning et al. identified 518 protein kinase genes and 106 protein kinase pseudogenes in human (Manning et al., 2002 Science). Most protein kinases contain a conserved catalytic domain belonging to the eukaryotic protein kinase (ePK) superfamily (all other protein kinases are classified as atypical protein kinases, or aPKs). ePKs are classified into 7 major groups, including AGC, CAMK, CK1, CMGC, STE, TK, and TKL, and are subdivided into families, and sometimes subfamilies, based on the sequence of their ePK domains. All 518 human kinases are listed as following, users can clikc into the URL link to investigate the detail information about the kinase and their substrates.

- Experimental protein kinases in human (Manning et al., 2002 Science)

No.KinaseGroupFamilySubfamilyGene SymbolDescription
1
AAK1OtherNAKBIKE AAK1 AP2 associated kinase 1 
2
ABL1TKAbl ABL1 v-abl Abelson murine leukemia viral oncogene homolog 1 
3
ABL2TKAbl ABL2 v-abl Abelson murine leukemia viral oncogene homolog 2 (arg, Abelson-related gene) 
4
ACKTKAck TNK2 tyrosine kinase, non-receptor, 2 
5
ACTR2TKLSTKRType2 ACVR2A activin A receptor, type IIA 
6
ACTR2BTKLSTKRType2 ACVR2B activin A receptor, type IIB 
7
ADCK1AtypicalABC1ABC1-B ADCK1 aarF domain containing kinase 1 
8
ADCK2AtypicalABC1ABC1-C ADCK2 aarF domain containing kinase 2 
9
ADCK3AtypicalABC1ABC1-A CABC1 chaperone, ABC1 activity of bc1 complex homolog (S. pombe) 
10
ADCK4AtypicalABC1ABC1-A ADCK4 aarF domain containing kinase 4 
11
ADCK5AtypicalABC1ABC1-B ADCK5 aarF domain containing kinase 5 
12
AKT1AGCAkt AKT1 v-akt murine thymoma viral oncogene homolog 1 
13
AKT2AGCAkt AKT2 v-akt murine thymoma viral oncogene homolog 2 
14
AKT3AGCAkt AKT3 v-akt murine thymoma viral oncogene homolog 3 (protein kinase B, gamma) 
15
ALKTKALK ALK anaplastic lymphoma kinase (Ki-1) 
16
ALK1TKLSTKRType1 ACVRL1 activin A receptor type II-like 1 
17
ALK2TKLSTKRType1 ACVR1 activin A receptor, type I 
18
ALK4TKLSTKRType1 ACVR1B activin A receptor, type IB 
19
ALK7TKLSTKRType1 ACVR1C activin A receptor, type IC 
20
AlphaK1AtypicalAlphaAlpha-Unclassified ALPK3 alpha-kinase 3 
21
AlphaK2AtypicalAlphaAlpha-Unclassified ALPK2 alpha-kinase 2 
22
AlphaK3AtypicalAlphaAlpha-Unclassified ALPK1 alpha-kinase 1 
23
AMPKa1CAMKCAMKLAMPK PRKAA1 protein kinase, AMP-activated, alpha 1 catalytic subunit 
24
AMPKa2CAMKCAMKLAMPK PRKAA2 protein kinase, AMP-activated, alpha 2 catalytic subunit 
25
ANKRD3TKLRIPK RIPK4 receptor-interacting serine-threonine kinase 4 
26
ANPaRGCRGC NPR1 natriuretic peptide receptor A/guanylate cyclase A (atrionatriuretic peptide receptor A) 
27
ANPbRGCRGC NPR2 natriuretic peptide receptor B/guanylate cyclase B (atrionatriuretic peptide receptor B) 
28
ARAFTKLRAFRAF ARAF v-raf murine sarcoma 3611 viral oncogene homolog 
29
ATMAtypicalPIKKATM ATM ataxia telangiectasia mutated (includes complementation groups A, C and D) 
30
ATRAtypicalPIKKATR ATR ataxia telangiectasia and Rad3 related 
31
AurAOtherAur AURKA aurora kinase A 
32
AurBOtherAur AURKB aurora kinase B 
33
AurCOtherAur AURKC aurora kinase C 
34
AXLTKAxl AXL AXL receptor tyrosine kinase 
35
BARK1AGCGRKBARK ADRBK1 adrenergic, beta, receptor kinase 1 
36
BARK2AGCGRKBARK ADRBK2 adrenergic, beta, receptor kinase 2 
37
BCKDKAtypicalPDHKBCKDK BCKDK branched chain ketoacid dehydrogenase kinase 
38
BCRAtypicalBCR BCR breakpoint cluster region 
39
BIKEOtherNAKBIKE BMP2K BMP2 inducible kinase 
40
BLKTKSrcSrcB BLK B lymphoid tyrosine kinase 
41
BMPR1ATKLSTKRType1 BMPR1A bone morphogenetic protein receptor, type IA 
42
BMPR1BTKLSTKRType1 BMPR1B bone morphogenetic protein receptor, type IB 
43
BMPR2TKLSTKRType2 BMPR2 bone morphogenetic protein receptor, type II (serine/threonine kinase) 
44
BMXTKTec BMX BMX non-receptor tyrosine kinase 
45
BRAFTKLRAFRAF BRAF v-raf murine sarcoma viral oncogene homolog B1 
46
BRD2AtypicalBRD BRD2 bromodomain containing 2 
47
BRD3AtypicalBRD BRD3 bromodomain containing 3 
48
BRD4AtypicalBRD BRD4 bromodomain containing 4 
49
BRDTAtypicalBRD BRDT bromodomain, testis-specific 
50
BRKTKSrcSRM PTK6 PTK6 protein tyrosine kinase 6 
51
BRSK1CAMKCAMKLBRSK BRSK1 BR serine/threonine kinase 1 
52
BRSK2CAMKCAMKLBRSK BRSK2 BR serine/threonine kinase 2 
53
BTKTKTec BTK Bruton agammaglobulinemia tyrosine kinase 
54
BUB1OtherBUB BUB1 BUB1 budding uninhibited by benzimidazoles 1 homolog (yeast) 
55
BUBR1OtherBUB BUB1B BUB1 budding uninhibited by benzimidazoles 1 homolog beta (yeast) 
56
CaMK1aCAMKCAMK1 CAMK1 calcium/calmodulin-dependent protein kinase I 
57
CaMK1bCAMKCAMK1 PNCK pregnancy upregulated non-ubiquitously expressed CaM kinase 
58
CaMK1dCAMKCAMK1 CAMK1D calcium/calmodulin-dependent protein kinase ID 
59
CaMK1gCAMKCAMK1 CAMK1G calcium/calmodulin-dependent protein kinase IG 
60
CaMK2aCAMKCAMK2 CAMK2A calcium/calmodulin-dependent protein kinase (CaM kinase) II alpha 
61
CaMK2bCAMKCAMK2 CAMK2B calcium/calmodulin-dependent protein kinase (CaM kinase) II beta 
62
CaMK2dCAMKCAMK2 CAMK2D calcium/calmodulin-dependent protein kinase (CaM kinase) II delta 
63
CaMK2gCAMKCAMK2 CAMK2G calcium/calmodulin-dependent protein kinase (CaM kinase) II gamma 
64
CaMK4CAMKCAMK1 CAMK4 calcium/calmodulin-dependent protein kinase IV 
65
CaMKK1OtherCAMKKMeta CAMKK1 calcium/calmodulin-dependent protein kinase kinase 1, alpha 
66
CaMKK2OtherCAMKKMeta CAMKK2 calcium/calmodulin-dependent protein kinase kinase 2, beta 
67
caMLCKCAMKMLCK MLCK MLCK protein 
68
CASKCAMKCASK CASK calcium/calmodulin-dependent serine protein kinase (MAGUK family) 
69
CCK4TKCCK4 PTK7 PTK7 protein tyrosine kinase 7 
70
CCRKCMGCCDKCCRK CCRK cell cycle related kinase 
71
CDC2CMGCCDKCDC2 CDC2 cell division cycle 2, G1 to S and G2 to M 
72
CDC7OtherCDC7 CDC7 cell division cycle 7 homolog (S. cerevisiae) 
73
CDK10CMGCCDKCDK10 CDK10 cyclin-dependent kinase (CDC2-like) 10 
74
CDK11CMGCCDKCDK8 CDC2L6 cell division cycle 2-like 6 (CDK8-like) 
75
CDK2CMGCCDKCDC2 CDK2 cyclin-dependent kinase 2 
76
CDK3CMGCCDKCDC2 CDK3 cyclin-dependent kinase 3 
77
CDK4CMGCCDKCDK4 CDK4 cyclin-dependent kinase 4 
78
CDK5CMGCCDKCDK5 CDK5 cyclin-dependent kinase 5 
79
CDK6CMGCCDKCDK4 CDK6 cyclin-dependent kinase 6 
80
CDK7CMGCCDKCDK7 CDK7 cyclin-dependent kinase 7 (MO15 homolog, Xenopus laevis, cdk-activating kinase) 
81
CDK8CMGCCDKCDK8 CDK8 cyclin-dependent kinase 8 
82
CDK9CMGCCDKCDK9 CDK9 cyclin-dependent kinase 9 (CDC2-related kinase) 
83
CDKL1CMGCCDKL CDKL1 cyclin-dependent kinase-like 1 (CDC2-related kinase) 
84
CDKL2CMGCCDKL CDKL2 cyclin-dependent kinase-like 2 (CDC2-related kinase) 
85
CDKL3CMGCCDKL CDKL3 cyclin-dependent kinase-like 3 
86
CDKL4CMGCCDKL CDKL4 cyclin-dependent kinase-like 4 
87
CDKL5CMGCCDKL CDKL5 cyclin-dependent kinase-like 5 
88
ChaK1AtypicalAlphaChaK TRPM7 transient receptor potential cation channel, subfamily M, member 7 
89
ChaK2AtypicalAlphaChaK TRPM6 transient receptor potential cation channel, subfamily M, member 6 
90
CHEDCMGCCDKCRK7 CDC2L5 cell division cycle 2-like 5 (cholinesterase-related cell division controller) 
91
CHK1CAMKCAMKLCHK1 CHEK1 CHK1 checkpoint homolog (S. pombe) 
92
CHK2CAMKRAD53 CHEK2 CHK2 checkpoint homolog (S. pombe) 
93
CK1aCK1CK1CK1-A CSNK1A1 casein kinase 1, alpha 1 
94
CK1a2CK1CK1CK1-A CSNK1A1L casein kinase 1, alpha 1-like 
95
CK1dCK1CK1CK1-D CSNK1D casein kinase 1, delta 
96
CK1eCK1CK1CK1-D CSNK1E casein kinase 1, epsilon 
97
CK1g1CK1CK1CK1-G CSNK1G1 casein kinase 1, gamma 1 
98
CK1g2CK1CK1CK1-G CSNK1G2 casein kinase 1, gamma 2 
99
CK1g3CK1CK1CK1-G CSNK1G3 casein kinase 1, gamma 3 
100
CK2a1CMGCCK2 CSNK2A1 casein kinase 2, alpha 1 polypeptide 
101
CK2a1-rsCMGCCK2   
102
CK2a2CMGCCK2 CSNK2A2 casein kinase 2, alpha prime polypeptide 
103
CLIK1OtherNKF4 STK35 serine/threonine kinase 35 
104
CLIK1LOtherNKF4 PDIK1L PDLIM1 interacting kinase 1 like 
105
CLK1CMGCCLK CLK1 CDC-like kinase 1 
106
CLK2CMGCCLK CLK2 CDC-like kinase 2 
107
CLK3CMGCCLK CLK3 CDC-like kinase 3 
108
CLK4CMGCCLK CLK4 CDC-like kinase 4 
109
COTSTESTE-Unique MAP3K8 mitogen-activated protein kinase kinase kinase 8 
110
CRIKAGCDMPKCRIK CIT citron (rho-interacting, serine/threonine kinase 21) 
111
CRK7CMGCCDKCRK7 CRKRS Cdc2-related kinase, arginine/serine-rich 
112
CSKTKCsk CSK c-src tyrosine kinase 
113
CTKTKCsk MATK megakaryocyte-associated tyrosine kinase 
114
CYGDRGCRGC GUCY2D guanylate cyclase 2D, membrane (retina-specific) 
115
CYGFRGCRGC GUCY2F guanylate cyclase 2F, retinal 
116
DAPK1CAMKDAPKDAPK DAPK1 death-associated protein kinase 1 
117
DAPK2CAMKDAPKDAPK DAPK2 death-associated protein kinase 2 
118
DAPK3CAMKDAPKDAPK DAPK3 death-associated protein kinase 3 
119
DCLK1CAMKDCAMKL DCAMKL1 doublecortin and CaM kinase-like 1 
120
DCLK2CAMKDCAMKL DCAMKL2 doublecortin and CaM kinase-like 2 
121
DCLK3CAMKDCAMKL DCAMKL3 doublecortin and CaM kinase-like 3 
122
DDR1TKDDR DDR1 discoidin domain receptor family, member 1 
123
DDR2TKDDR DDR2 discoidin domain receptor family, member 2 
124
DLKTKLMLKLZK MAP3K12 mitogen-activated protein kinase kinase kinase 12 
125
DMPK1AGCDMPKGEK DMPK dystrophia myotonica-protein kinase 
126
DMPK2AGCDMPKGEK CDC42BPG CDC42 binding protein kinase gamma (DMPK-like) 
127
DNAPKAtypicalPIKKDNAPK PRKDC protein kinase, DNA-activated, catalytic polypeptide 
128
DRAK1CAMKDAPKDRAK STK17A serine/threonine kinase 17a (apoptosis-inducing) 
129
DRAK2CAMKDAPKDRAK STK17B serine/threonine kinase 17b (apoptosis-inducing) 
130
DYRK1ACMGCDYRKDYRK1 DYRK1A dual-specificity tyrosine-(Y)-phosphorylation regulated kinase 1A 
131
DYRK1BCMGCDYRKDYRK1 DYRK1B dual-specificity tyrosine-(Y)-phosphorylation regulated kinase 1B 
132
DYRK2CMGCDYRKDYRK2 DYRK2 dual-specificity tyrosine-(Y)-phosphorylation regulated kinase 2 
133
DYRK3CMGCDYRKDYRK2 DYRK3 dual-specificity tyrosine-(Y)-phosphorylation regulated kinase 3 
134
DYRK4CMGCDYRKDYRK2 DYRK4 dual-specificity tyrosine-(Y)-phosphorylation regulated kinase 4 
135
eEF2KAtypicalAlphaeEF2K EEF2K eukaryotic elongation factor-2 kinase 
136
EGFRTKEGFR EGFR epidermal growth factor receptor (erythroblastic leukemia viral (v-erb-b) oncogene homolog, avian) 
137
EphA1TKEph EPHA1 EPH receptor A1 
138
EphA10TKEph EPHA10 EPH receptor A10 
139
EphA2TKEph EPHA2 EPH receptor A2 
140
EphA3TKEph EPHA3 EPH receptor A3 
141
EphA4TKEph EPHA4 EPH receptor A4 
142
EphA5TKEph EPHA5 EPH receptor A5 
143
EphA6TKEph EPHA6 EPH receptor A6 
144
EphA7TKEph EPHA7 EPH receptor A7 
145
EphA8TKEph EPHA8 EPH receptor A8 
146
EphB1TKEph EPHB1 EPH receptor B1 
147
EphB2TKEph EPHB2 EPH receptor B2 
148
EphB3TKEph EPHB3 EPH receptor B3 
149
EphB4TKEph EPHB4 EPH receptor B4 
150
EphB6TKEph EPHB6 EPH receptor B6 
151
ErbB2TKEGFR ERBB2 v-erb-b2 erythroblastic leukemia viral oncogene homolog 2, neuro/glioblastoma derived oncogene homolog (avian) 
152
ErbB3TKEGFR ERBB3 v-erb-b2 erythroblastic leukemia viral oncogene homolog 3 (avian) 
153
ErbB4TKEGFR ERBB4 v-erb-a erythroblastic leukemia viral oncogene homolog 4 (avian) 
154
Erk1CMGCMAPKERK1 MAPK3 mitogen-activated protein kinase 3 
155
Erk2CMGCMAPKERK1 MAPK1 mitogen-activated protein kinase 1 
156
Erk3CMGCMAPKERK3 MAPK6 mitogen-activated protein kinase 6 
157
Erk4CMGCMAPKERK3 MAPK4 mitogen-activated protein kinase 4 
158
Erk5CMGCMAPKERK5 MAPK7 mitogen-activated protein kinase 7 
159
Erk7CMGCMAPKERK7 MAPK15 mitogen-activated protein kinase 15 
160
FAKTKFAK PTK2 PTK2 protein tyrosine kinase 2 
161
FASTKAtypicalFAST FASTK Fas-activated serine/threonine kinase 
162
FERTKFer FER fer (fps/fes related) tyrosine kinase (phosphoprotein NCP94) 
163
FESTKFer FES feline sarcoma oncogene 
164
FGFR1TKFGFR FGFR1 fibroblast growth factor receptor 1 (fms-related tyrosine kinase 2, Pfeiffer syndrome) 
165
FGFR2TKFGFR FGFR2 fibroblast growth factor receptor 2 (bacteria-expressed kinase, keratinocyte growth factor receptor, craniofacial dysostosis 1, Crouzon syndrome, Pfeiffer syndrome, Jackson-Weiss syndrome) 
166
FGFR3TKFGFR FGFR3 fibroblast growth factor receptor 3 (achondroplasia, thanatophoric dwarfism) 
167
FGFR4TKFGFR FGFR4 fibroblast growth factor receptor 4 
168
FGRTKSrcSrcA FGR Gardner-Rasheed feline sarcoma viral (v-fgr) oncogene homolog 
169
FLT1TKVEGFR FLT1 fms-related tyrosine kinase 1 (vascular endothelial growth factor/vascular permeability factor receptor) 
170
FLT3TKPDGFR FLT3 fms-related tyrosine kinase 3 
171
FLT4TKVEGFR FLT4 fms-related tyrosine kinase 4 
172
FMSTKPDGFR CSF1R colony stimulating factor 1 receptor, formerly McDonough feline sarcoma viral (v-fms) oncogene homolog 
173
FRAPAtypicalPIKKFRAP FRAP1 FK506 binding protein 12-rapamycin associated protein 1 
174
FRKTKSrcFrk FRK fyn-related kinase 
175
FusedOtherULKFused STK36 serine/threonine kinase 36, fused homolog (Drosophila) 
176
FYNTKSrcSrcA FYN FYN oncogene related to SRC, FGR, YES 
177
G11AtypicalG11 STK19 serine/threonine kinase 19 
178
GAKOtherNAKGAK GAK cyclin G associated kinase 
179
GCKSTESTE20KHS MAP4K2 mitogen-activated protein kinase kinase kinase kinase 2 
180
GCN2OtherPEKGCN2 EIF2AK4 eukaryotic translation initiation factor 2 alpha kinase 4 
181
GPRK4AGCGRKGRK GRK4 G protein-coupled receptor kinase 4 
182
GPRK5AGCGRKGRK GRK5 G protein-coupled receptor kinase 5 
183
GPRK6AGCGRKGRK GRK6 G protein-coupled receptor kinase 6 
184
GPRK7AGCGRKGRK GRK7 G protein-coupled receptor kinase 7 
185
GSK3ACMGCGSK GSK3A glycogen synthase kinase 3 alpha 
186
GSK3BCMGCGSK GSK3B glycogen synthase kinase 3 beta 
187
H11AtypicalH11 HSPB8 heat shock 22kDa protein 8 
188
HaspinOtherHaspin GSG2 germ cell associated 2 (haspin) 
189
HCKTKSrcSrcB HCK hemopoietic cell kinase 
190
HGKSTESTE20MSN MAP4K4 mitogen-activated protein kinase kinase kinase kinase 4 
191
HH498TKLMLKHH498 TNNI3K TNNI3 interacting kinase 
192
HIPK1CMGCDYRKHIPK HIPK1 homeodomain interacting protein kinase 1 
193
HIPK2CMGCDYRKHIPK HIPK2 homeodomain interacting protein kinase 2 
194
HIPK3CMGCDYRKHIPK HIPK3 homeodomain interacting protein kinase 3 
195
HIPK4CMGCDYRKHIPK HIPK4 homeodomain interacting protein kinase 4 
196
HPK1STESTE20KHS MAP4K1 mitogen-activated protein kinase kinase kinase kinase 1 
197
HRIOtherPEKHRI EIF2AK1 eukaryotic translation initiation factor 2-alpha kinase 1 
198
HSERRGCRGC GUCY2C guanylate cyclase 2C (heat stable enterotoxin receptor) 
199
HUNKCAMKCAMKLHUNK HUNK hormonally upregulated Neu-associated kinase 
200
ICKCMGCRCK ICK intestinal cell (MAK-like) kinase 
201
IGF1RTKInsR IGF1R insulin-like growth factor 1 receptor 
202
IKKaOtherIKK CHUK conserved helix-loop-helix ubiquitous kinase 
203
IKKbOtherIKK IKBKB inhibitor of kappa light polypeptide gene enhancer in B-cells, kinase beta 
204
IKKeOtherIKK IKBKE inhibitor of kappa light polypeptide gene enhancer in B-cells, kinase epsilon 
205
ILKTKLMLKILK ILK integrin-linked kinase 
206
INSRTKInsR INSR insulin receptor 
207
IRAK1TKLIRAK IRAK1 interleukin-1 receptor-associated kinase 1 
208
IRAK2TKLIRAK IRAK2 interleukin-1 receptor-associated kinase 2 
209
IRAK3TKLIRAK IRAK3 interleukin-1 receptor-associated kinase 3 
210
IRAK4TKLIRAK IRAK4 interleukin-1 receptor-associated kinase 4 
211
IRE1OtherIRE ERN1 endoplasmic reticulum to nucleus signalling 1 
212
IRE2OtherIRE ERN2 endoplasmic reticulum to nucleus signalling 2 
213
IRRTKInsR INSRR insulin receptor-related receptor 
214
ITKTKTec ITK IL2-inducible T-cell kinase 
215
JAK1TKJakA JAK1 Janus kinase 1 (a protein tyrosine kinase) 
216
JAK2TKJakA JAK2 Janus kinase 2 (a protein tyrosine kinase) 
217
JAK3TKJakA JAK3 Janus kinase 3 (a protein tyrosine kinase, leukocyte) 
218
JNK1CMGCMAPKJNK MAPK8 mitogen-activated protein kinase 8 
219
JNK2CMGCMAPKJNK MAPK9 mitogen-activated protein kinase 9 
220
JNK3CMGCMAPKJNK MAPK10 mitogen-activated protein kinase 10 
221
KDRTKVEGFR KDR kinase insert domain receptor (a type III receptor tyrosine kinase) 
222
KHS1STESTE20KHS MAP4K5 mitogen-activated protein kinase kinase kinase kinase 5 
223
KHS2STESTE20KHS MAP4K3 mitogen-activated protein kinase kinase kinase kinase 3 
224
KISOtherKIS UHMK1 U2AF homology motif (UHM) kinase 1 
225
KITTKPDGFR KIT v-kit Hardy-Zuckerman 4 feline sarcoma viral oncogene homolog 
226
KSR1TKLRAFKSR KSR1 kinase suppressor of ras 1 
227
KSR2TKLRAFKSR KSR2 kinase suppressor of ras 2 
228
LATS1AGCNDR LATS1 LATS, large tumor suppressor, homolog 1 (Drosophila) 
229
LATS2AGCNDR LATS2 LATS, large tumor suppressor, homolog 2 (Drosophila) 
230
LCKTKSrcSrcB LCK lymphocyte-specific protein tyrosine kinase 
231
LIMK1TKLLISKLIMK LIMK1 LIM domain kinase 1 
232
LIMK2TKLLISKLIMK LIMK2 LIM domain kinase 2 
233
LKB1CAMKCAMKLLKB STK11 serine/threonine kinase 11 
234
LMR1TKLmr AATK apoptosis-associated tyrosine kinase 
235
LMR2TKLmr LMTK2 lemur tyrosine kinase 2 
236
LMR3TKLmr LMTK3 lemur tyrosine kinase 3 
237
LOKSTESTE20SLK STK10 serine/threonine kinase 10 
238
LRRK1TKLLRRK LRRK1 leucine-rich repeat kinase 1 
239
LRRK2TKLLRRK LRRK2 leucine-rich repeat kinase 2 
240
LTKTKALK LTK leukocyte tyrosine kinase 
241
LYNTKSrcSrcB LYN v-yes-1 Yamaguchi sarcoma viral related oncogene homolog 
242
LZKTKLMLKLZK MAP3K13 mitogen-activated protein kinase kinase kinase 13 
243
MAKCMGCRCK MAK male germ cell-associated kinase 
244
MAP2K1STESTE7MEK1 MAP2K1 mitogen-activated protein kinase kinase 1 
245
MAP2K2STESTE7MEK1 MAP2K2 mitogen-activated protein kinase kinase 2 
246
MAP2K3STESTE7MEK3 MAP2K3 mitogen-activated protein kinase kinase 3 
247
MAP2K4STESTE7MEK4 MAP2K4 mitogen-activated protein kinase kinase 4 
248
MAP2K5STESTE7MEK5 MAP2K5 mitogen-activated protein kinase kinase 5 
249
MAP2K6STESTE7MEK3 MAP2K6 mitogen-activated protein kinase kinase 6 
250
MAP2K7STESTE7MEK7 MAP2K7 mitogen-activated protein kinase kinase 7 
251
MAP3K1STESTE11MEKK1 MAP3K1 mitogen-activated protein kinase kinase kinase 1 
252
MAP3K2STESTE11MEKK2 MAP3K2 mitogen-activated protein kinase kinase kinase 2 
253
MAP3K3STESTE11MEKK2 MAP3K3 mitogen-activated protein kinase kinase kinase 3 
254
MAP3K4STESTE11MEKK4 MAP3K4 mitogen-activated protein kinase kinase kinase 4 
255
MAP3K5STESTE11ASK MAP3K5 mitogen-activated protein kinase kinase kinase 5 
256
MAP3K6STESTE11ASK MAP3K6 mitogen-activated protein kinase kinase kinase 6 
257
MAP3K7STESTE11ASK MAP3K15 mitogen-activated protein kinase kinase kinase 15 
258
MAP3K8STESTE11MEKK15 YSK4 yeast Sps1/Ste20-related kinase 4 (S. cerevisiae) 
259
MAPKAPK2CAMKMAPKAPKMAPKAPK MAPKAPK2 mitogen-activated protein kinase-activated protein kinase 2 
260
MAPKAPK3CAMKMAPKAPKMAPKAPK MAPKAPK3 mitogen-activated protein kinase-activated protein kinase 3 
261
MAPKAPK5CAMKMAPKAPKMAPKAPK MAPKAPK5 mitogen-activated protein kinase-activated protein kinase 5 
262
MARK1CAMKCAMKLMARK MARK1 MAP/microtubule affinity-regulating kinase 1 
263
MARK2CAMKCAMKLMARK MARK2 MAP/microtubule affinity-regulating kinase 2 
264
MARK3CAMKCAMKLMARK MARK3 MAP/microtubule affinity-regulating kinase 3 
265
MARK4CAMKCAMKLMARK MARK4 MAP/microtubule affinity-regulating kinase 4 
266
MAST1AGCMASTMAST MAST1 microtubule associated serine/threonine kinase 1 
267
MAST2AGCMASTMAST MAST2 microtubule associated serine/threonine kinase 2 
268
MAST3AGCMASTMAST MAST3 microtubule associated serine/threonine kinase 3 
269
MAST4AGCMASTMAST LOC375449 similar to microtubule associated testis specific serine/threonine protein kinase 
270
MASTLAGCMASTMASTL MASTL microtubule associated serine/threonine kinase-like 
271
MELKCAMKCAMKLMELK MELK maternal embryonic leucine zipper kinase 
272
MERTKAxl MERTK c-mer proto-oncogene tyrosine kinase 
273
METTKMet MET met proto-oncogene (hepatocyte growth factor receptor) 
274
MINKSTESTE20MSN MINK1 misshapen-like kinase 1 (zebrafish) 
275
MISR2TKLSTKRType2 AMHR2 anti-Mullerian hormone receptor, type II 
276
MLK1TKLMLKMLK MAP3K9 mitogen-activated protein kinase kinase kinase 9 
277
MLK2TKLMLKMLK MAP3K10 mitogen-activated protein kinase kinase kinase 10 
278
MLK3TKLMLKMLK MAP3K11 mitogen-activated protein kinase kinase kinase 11 
279
MLK4TKLMLKMLK KIAA1804 mixed lineage kinase 4 
280
MLKLTKLTKL-Unique MLKL mixed lineage kinase domain-like 
281
MNK1CAMKMAPKAPKMNK MKNK1 MAP kinase interacting serine/threonine kinase 1 
282
MNK2CAMKMAPKAPKMNK MKNK2 MAP kinase interacting serine/threonine kinase 2 
283
MOKCMGCRCK RAGE renal tumor antigen 
284
MOSOtherMOS MOS v-mos Moloney murine sarcoma viral oncogene homolog 
285
MPSK1OtherNAKMPSK STK16 serine/threonine kinase 16 
286
MRCKaAGCDMPKGEK CDC42BPA CDC42 binding protein kinase alpha (DMPK-like) 
287
MRCKbAGCDMPKGEK CDC42BPB CDC42 binding protein kinase beta (DMPK-like) 
288
MSK1AGCRSKMSK RPS6KA5 ribosomal protein S6 kinase, 90kDa, polypeptide 5 
289
MSK2AGCRSKMSK RPS6KA4 ribosomal protein S6 kinase, 90kDa, polypeptide 4 
290
MSSK1CMGCSRPK SRPK3 SFRS protein kinase 3 
291
MST1STESTE20MST STK4 serine/threonine kinase 4 
292
MST2STESTE20MST STK3 serine/threonine kinase 3 (STE20 homolog, yeast) 
293
MST3STESTE20YSK STK24 serine/threonine kinase 24 (STE20 homolog, yeast) 
294
MST4STESTE20YSK RP6-213H19.1 serine/threonine protein kinase MST4 
295
MUSKTKMusk MUSK muscle, skeletal, receptor tyrosine kinase 
296
MYO3ASTESTE20NinaC MYO3A myosin IIIA 
297
MYO3BSTESTE20NinaC MYO3B myosin IIIB 
298
MYT1OtherWEE PKMYT1 protein kinase, membrane associated tyrosine/threonine 1 
299
NDR1AGCNDR STK38 serine/threonine kinase 38 
300
NDR2AGCNDR STK38L serine/threonine kinase 38 like 
301
NEK1OtherNEKNEK1 NEK1 NIMA (never in mitosis gene a)-related kinase 1 
302
NEK10OtherNEKNEK10 NEK10 NIMA (never in mitosis gene a)- related kinase 10 
303
NEK11OtherNEKNEK11 NEK11 NIMA (never in mitosis gene a)- related kinase 11 
304
NEK2OtherNEKNEK2 NEK2 NIMA (never in mitosis gene a)-related kinase 2 
305
NEK3OtherNEKNEK1 NEK3 NIMA (never in mitosis gene a)-related kinase 3 
306
NEK4OtherNEKNEK4 NEK4 NIMA (never in mitosis gene a)-related kinase 4 
307
NEK5OtherNEKNEK1 NEK5 NIMA (never in mitosis gene a)-related kinase 5 
308
NEK6OtherNEKNEK6 NEK6 NIMA (never in mitosis gene a)-related kinase 6 
309
NEK7OtherNEKNEK6 NEK7 NIMA (never in mitosis gene a)-related kinase 7 
310
NEK8OtherNEKNEK8 NEK8 NIMA (never in mitosis gene a)- related kinase 8 
311
NEK9OtherNEKNEK8 NEK9 NIMA (never in mitosis gene a)- related kinase 9 
312
NIKSTESTE-Unique MAP3K14 mitogen-activated protein kinase kinase kinase 14 
313
NIM1CAMKCAMKLNIM1 MGC42105 hypothetical protein MGC42105 
314
NLKCMGCMAPKnmo NLK nemo-like kinase 
315
NRBP1OtherNRBP NRBP1 nuclear receptor binding protein 1 
316
NRBP2OtherNRBP NRBP2 nuclear receptor binding protein 2 
317
NRKSTESTE20MSN NRK Nik related kinase 
318
NuaK1CAMKCAMKLNuaK NUAK1 NUAK family, SNF1-like kinase, 1 
319
NuaK2CAMKCAMKLNuaK NUAK2 NUAK family, SNF1-like kinase, 2 
320
ObscnCAMKTrio OBSCN obscurin, cytoskeletal calmodulin and titin-interacting RhoGEF 
321
OSR1STESTE20FRAY OXSR1 oxidative-stress responsive 1 
322
p38aCMGCMAPKp38 MAPK14 mitogen-activated protein kinase 14 
323
p38bCMGCMAPKp38 MAPK11 mitogen-activated protein kinase 11 
324
p38dCMGCMAPKp38 MAPK13 mitogen-activated protein kinase 13 
325
p38gCMGCMAPKp38 MAPK12 mitogen-activated protein kinase 12 
326
p70S6KAGCRSKp70 RPS6KB1 ribosomal protein S6 kinase, 70kDa, polypeptide 1 
327
p70S6KbAGCRSKp70 RPS6KB2 ribosomal protein S6 kinase, 70kDa, polypeptide 2 
328
PAK1STESTE20PAKA PAK1 p21/Cdc42/Rac1-activated kinase 1 (STE20 homolog, yeast) 
329
PAK2STESTE20PAKA PAK2 p21 (CDKN1A)-activated kinase 2 
330
PAK3STESTE20PAKA PAK3 p21 (CDKN1A)-activated kinase 3 
331
PAK4STESTE20PAKB PAK4 p21(CDKN1A)-activated kinase 4 
332
PAK5STESTE20PAKB PAK7 p21(CDKN1A)-activated kinase 7 
333
PAK6STESTE20PAKB PAK6 p21(CDKN1A)-activated kinase 6 
334
PASKCAMKCAMKLPASK PASK PAS domain containing serine/threonine kinase 
335
PBKOtherTOPK PBK PDZ binding kinase 
336
PCTAIRE1CMGCCDKPCTAIRE PCTK1 PCTAIRE protein kinase 1 
337
PCTAIRE2CMGCCDKPCTAIRE PCTK2 PCTAIRE protein kinase 2 
338
PCTAIRE3CMGCCDKPCTAIRE PCTK3 PCTAIRE protein kinase 3 
339
PDGFRaTKPDGFR PDGFRA platelet-derived growth factor receptor, alpha polypeptide 
340
PDGFRbTKPDGFR PDGFRB platelet-derived growth factor receptor, beta polypeptide 
341
PDHK1AtypicalPDHKPDHK PDK1 pyruvate dehydrogenase kinase, isozyme 1 
342
PDHK2AtypicalPDHKPDHK PDK2 pyruvate dehydrogenase kinase, isozyme 2 
343
PDHK3AtypicalPDHKPDHK PDK3 pyruvate dehydrogenase kinase, isozyme 3 
344
PDHK4AtypicalPDHKPDHK PDK4 pyruvate dehydrogenase kinase, isozyme 4 
345
PDK1AGCPDK1 PDPK1 3-phosphoinositide dependent protein kinase-1 
346
PEKOtherPEKPEK EIF2AK3 eukaryotic translation initiation factor 2-alpha kinase 3 
347
PFTAIRE1CMGCCDKPFTAIRE PFTK1 PFTAIRE protein kinase 1 
348
PFTAIRE2CMGCCDKPFTAIRE ALS2CR7 amyotrophic lateral sclerosis 2 (juvenile) chromosome region, candidate 7 
349
PHKg1CAMKPHK PHKG1 phosphorylase kinase, gamma 1 (muscle) 
350
PHKg2CAMKPHK PHKG2 phosphorylase kinase, gamma 2 (testis) 
351
PIK3R4OtherVPS15 PIK3R4 phosphoinositide-3-kinase, regulatory subunit 4, p150 
352
PIM1CAMKPIM PIM1 pim-1 oncogene 
353
PIM2CAMKPIM PIM2 pim-2 oncogene 
354
PIM3CAMKPIM PIM3 pim-3 oncogene 
355
PINK1OtherNKF2 PINK1 PTEN induced putative kinase 1 
356
PITSLRECMGCCDKPITSLRE CDC2L2 cell division cycle 2-like 2 (PITSLRE proteins) 
357
PKACaAGCPKA PRKACA protein kinase, cAMP-dependent, catalytic, alpha 
358
PKACbAGCPKA PRKACB protein kinase, cAMP-dependent, catalytic, beta 
359
PKACgAGCPKA PRKACG protein kinase, cAMP-dependent, catalytic, gamma 
360
PKCaAGCPKCAlpha PRKCA protein kinase C, alpha 
361
PKCbAGCPKCAlpha PRKCB1 protein kinase C, beta 1 
362
PKCdAGCPKCDelta PRKCD protein kinase C, delta 
363
PKCeAGCPKCEta PRKCE protein kinase C, epsilon 
364
PKCgAGCPKCAlpha PRKCG protein kinase C, gamma 
365
PKChAGCPKCEta PRKCH protein kinase C, eta 
366
PKCiAGCPKCIota PRKCI protein kinase C, iota 
367
PKCtAGCPKCDelta PRKCQ protein kinase C, theta 
368
PKCzAGCPKCIota PRKCZ protein kinase C, zeta 
369
PKD1CAMKPKD PRKD1 protein kinase D1 
370
PKD2CAMKPKD PRKD2 protein kinase D2 
371
PKD3CAMKPKD PRKD3 protein kinase D3 
372
PKG1AGCPKG PRKG1 protein kinase, cGMP-dependent, type I 
373
PKG2AGCPKG PRKG2 protein kinase, cGMP-dependent, type II 
374
PKN1AGCPKN PKN1 protein kinase N1 
375
PKN2AGCPKN PKN2 protein kinase N2 
376
PKN3AGCPKN PKN3 protein kinase N3 
377
PKROtherPEKPKR EIF2AK2 eukaryotic translation initiation factor 2-alpha kinase 2 
378
PLK1OtherPLKPLK1 PLK1 polo-like kinase 1 (Drosophila) 
379
PLK2OtherPLKPLK2 PLK2 polo-like kinase 2 (Drosophila) 
380
PLK3OtherPLKPLK2 PLK3 polo-like kinase 3 (Drosophila) 
381
PLK4OtherPLKSAK PLK4 polo-like kinase 4 (Drosophila) 
382
PRKXAGCPKA PRKX protein kinase, X-linked 
383
PRKYAGCPKA PRKY protein kinase, Y-linked 
384
PRP4CMGCDYRKPRP4 PRPF4B PRP4 pre-mRNA processing factor 4 homolog B (yeast) 
385
PRPKOtherBud32 TP53RK TP53 regulating kinase 
386
PSKH1CAMKPSK PSKH1 protein serine kinase H1 
387
PSKH2CAMKPSK PSKH2 protein serine kinase H2 
388
PYK2TKFAK PTK2B PTK2B protein tyrosine kinase 2 beta 
389
QIKCAMKCAMKLQIK SNF1LK2 SNF1-like kinase 2 
390
QSKCAMKCAMKLQIK KIAA0999 KIAA0999 protein 
391
RAF1TKLRAFRAF RAF1 v-raf-1 murine leukemia viral oncogene homolog 1 
392
RETTKRet RET ret proto-oncogene (multiple endocrine neoplasia and medullary thyroid carcinoma 1, Hirschsprung disease) 
393
RHOKAGCGRKGRK GRK1 G protein-coupled receptor kinase 1 
394
RIOK1AtypicalRIORIO1 RIOK1 RIO kinase 1 (yeast) 
395
RIOK2AtypicalRIORIO2 RIOK2 RIO kinase 2 (yeast) 
396
RIOK3AtypicalRIORIO3 RIOK3 RIO kinase 3 (yeast) 
397
RIPK1TKLRIPK RIPK1 receptor (TNFRSF)-interacting serine-threonine kinase 1 
398
RIPK2TKLRIPK RIPK2 receptor-interacting serine-threonine kinase 2 
399
RIPK3TKLRIPK RIPK3 receptor-interacting serine-threonine kinase 3 
400
RNAseLOtherOther-Unique RNASEL ribonuclease L (2',5'-oligoisoadenylate synthetase-dependent) 
401
ROCK1AGCDMPKROCK ROCK1 Rho-associated, coiled-coil containing protein kinase 1 
402
ROCK2AGCDMPKROCK ROCK2 Rho-associated, coiled-coil containing protein kinase 2 
403
RONTKMet MST1R macrophage stimulating 1 receptor (c-met-related tyrosine kinase) 
404
ROR1TKRor ROR1 receptor tyrosine kinase-like orphan receptor 1 
405
ROR2TKRor ROR2 receptor tyrosine kinase-like orphan receptor 2 
406
ROSTKSev ROS1 v-ros UR2 sarcoma virus oncogene homolog 1 (avian) 
407
RSK1AGCRSKRSK RPS6KA2 ribosomal protein S6 kinase, 90kDa, polypeptide 2 
408
RSK2AGCRSKRSK RPS6KA3 ribosomal protein S6 kinase, 90kDa, polypeptide 3 
409
RSK3AGCRSKRSK RPS6KA1 ribosomal protein S6 kinase, 90kDa, polypeptide 1 
410
RSK4AGCRSKRSK RPS6KA6 ribosomal protein S6 kinase, 90kDa, polypeptide 6 
411
RSKL1AGCRSKL RPS6KC1 ribosomal protein S6 kinase, 52kDa, polypeptide 1 
412
RSKL2AGCRSKL RPS6KL1 ribosomal protein S6 kinase-like 1 
413
RYKTKRyk RYK RYK receptor-like tyrosine kinase 
414
SBKOtherNKF1 SBK1 SH3-binding domain kinase 1 
415
SCYL1OtherSCY1 SCYL1 SCY1-like 1 (S. cerevisiae) 
416
SCYL2OtherSCY1 SCYL2 SCY1-like 2 (S. cerevisiae) 
417
SCYL3OtherSCY1 SCYL3 SCY1-like 3 (S. cerevisiae) 
418
SgK069OtherNKF1 LOC646643 similar to protein kinase Bsk146 
419
SgK071OtherSgK071 C9orf96 chromosome 9 open reading frame 96 
420
SgK085CAMKMLCK RP11-145H9.1 hypothetical protein LOC340156 
421
SGK1AGCSGK SGK serum/glucocorticoid regulated kinase 
422
SgK110OtherNKF1   
423
SgK196OtherOther-Unique FLJ23356 hypothetical protein FLJ23356 
424
SGK2AGCSGK SGK2 serum/glucocorticoid regulated kinase 2 
425
SgK223OtherNKF3 DKFZp761P0423 homolog of rat pragma of Rnd2 
426
SgK269OtherNKF3 KIAA2002 KIAA2002 protein 
427
SgK288TKLRIPK ANKK1 ankyrin repeat and kinase domain containing 1 
428
SGK3AGCSGK SGK3 serum/glucocorticoid regulated kinase family, member 3 
429
SgK307OtherNKF5 TEX14 testis expressed sequence 14 
430
SgK396OtherOther-Unique STK31 serine/threonine kinase 31 
431
SgK424OtherNKF5   
432
SgK493OtherSgK493 LOC91461 hypothetical protein BC007901 
433
SgK494AGCRSKR FLJ25006 hypothetical protein FLJ25006 
434
SgK495CAMKSgK495 STK40 serine/threonine kinase 40 
435
SgK496OtherSgK496 RIPK5 receptor interacting protein kinase 5 
436
SIKCAMKCAMKLQIK SNF1LK SNF1-like kinase 
437
skMLCKCAMKMLCK MYLK2 myosin light chain kinase 2, skeletal muscle 
438
SLKSTESTE20SLK SLK STE20-like kinase (yeast) 
439
SlobOtherSlob PXK PX domain containing serine/threonine kinase 
440
SMG1AtypicalPIKKSMG1 SMG1 PI-3-kinase-related kinase SMG-1 
441
smMLCKCAMKMLCK MYLK myosin, light chain kinase 
442
SNRKCAMKCAMKLSNRK SNRK SNF related kinase 
443
SPEGCAMKTrio LOC729871 similar to aortic preferentially expressed gene 1 
444
SRCTKSrcSrcA SRC v-src sarcoma (Schmidt-Ruppin A-2) viral oncogene homolog (avian) 
445
SRMTKSrcSRM SRMS src-related kinase lacking C-terminal regulatory tyrosine and N-terminal myristylation sites 
446
SRPK1CMGCSRPK SRPK1 SFRS protein kinase 1 
447
SRPK2CMGCSRPK SRPK2 SFRS protein kinase 2 
448
SSTKCAMKTSSK TSSK6 testis-specific serine kinase 6 
449
STK33CAMKCAMK-Unique STK33 serine/threonine kinase 33 
450
STLK3STESTE20FRAY STK39 serine threonine kinase 39 (STE20/SPS1 homolog, yeast) 
451
STLK5STESTE20STLK LYK5 protein kinase LYK5 
452
STLK6STESTE20STLK ALS2CR2 amyotrophic lateral sclerosis 2 (juvenile) chromosome region, candidate 2 
453
STLK6-rsSTESTE20STLK   
454
SuRTK106TKTK-Unique STYK1 serine/threonine/tyrosine kinase 1 
455
SYKTKSyk SYK spleen tyrosine kinase 
456
TAF1AtypicalTAF1 TAF1 TAF1 RNA polymerase II, TATA box binding protein (TBP)-associated factor, 250kDa 
457
TAF1LAtypicalTAF1 TAF1L TAF1-like RNA polymerase II, TATA box binding protein (TBP)-associated factor, 210kDa 
458
TAK1TKLMLKTAK1 MAP3K7 mitogen-activated protein kinase kinase kinase 7 
459
TAO1STESTE20TAO TAOK1 TAO kinase 1 
460
TAO2STESTE20TAO TAOK2 TAO kinase 2 
461
TAO3STESTE20TAO TAOK3 TAO kinase 3 
462
TBCKOtherTBCK MGC16169 hypothetical protein MGC16169 
463
TBK1OtherIKK TBK1 TANK-binding kinase 1 
464
TECTKTec TEC tec protein tyrosine kinase 
465
TESK1TKLLISKTESK TESK1 testis-specific kinase 1 
466
TESK2TKLLISKTESK TESK2 testis-specific kinase 2 
467
TGFbR1TKLSTKRType1 TGFBR1 transforming growth factor, beta receptor I (activin A receptor type II-like kinase, 53kDa) 
468
TGFbR2TKLSTKRType2 TGFBR2 transforming growth factor, beta receptor II (70/80kDa) 
469
TIE1TKTie TIE1 tyrosine kinase with immunoglobulin-like and EGF-like domains 1 
470
TIE2TKTie TEK TEK tyrosine kinase, endothelial (venous malformations, multiple cutaneous and mucosal) 
471
TIF1aAtypicalTIF1 TRIM24 tripartite motif-containing 24 
472
TIF1bAtypicalTIF1 TRIM28 tripartite motif-containing 28 
473
TIF1gAtypicalTIF1 TRIM33 tripartite motif-containing 33 
474
TLK1OtherTLK TLK1 tousled-like kinase 1 
475
TLK2OtherTLK TLK2 tousled-like kinase 2 
476
TNIKSTESTE20MSN TNIK TRAF2 and NCK interacting kinase 
477
TNK1TKAck TNK1 tyrosine kinase, non-receptor, 1 
478
TradCAMKTrio KALRN kalirin, RhoGEF kinase 
479
Trb1CAMKTrbl TRIB1 tribbles homolog 1 (Drosophila) 
480
Trb2CAMKTrbl TRIB2 tribbles homolog 2 (Drosophila) 
481
Trb3CAMKTrbl TRIB3 tribbles homolog 3 (Drosophila) 
482
TrioCAMKTrio TRIO triple functional domain (PTPRF interacting) 
483
TRKATKTrk NTRK1 neurotrophic tyrosine kinase, receptor, type 1 
484
TRKBTKTrk NTRK2 neurotrophic tyrosine kinase, receptor, type 2 
485
TRKCTKTrk NTRK3 neurotrophic tyrosine kinase, receptor, type 3 
486
TRRAPAtypicalPIKKTRRAP TRRAP transformation/transcription domain-associated protein 
487
TSSK1CAMKTSSK TSSK1B testis-specific serine kinase 1B 
488
TSSK2CAMKTSSK TSSK2 testis-specific serine kinase 2 
489
TSSK3CAMKTSSK TSSK3 testis-specific serine kinase 3 
490
TSSK4CAMKTSSK TSSK4 testis-specific serine kinase 4 
491
TTBK1CK1TTBK TTBK1 tau tubulin kinase 1 
492
TTBK2CK1TTBK TTBK2 tau tubulin kinase 2 
493
TTKOtherTTK TTK TTK protein kinase 
494
TTNCAMKMLCK TTN titin 
495
TXKTKTec TXK TXK tyrosine kinase 
496
TYK2TKJakA TYK2 tyrosine kinase 2 
497
TYRO3TKAxl TYRO3 TYRO3 protein tyrosine kinase 
498
ULK1OtherULKULK ULK1 unc-51-like kinase 1 (C. elegans) 
499
ULK2OtherULKULK ULK2 unc-51-like kinase 2 (C. elegans) 
500
ULK3OtherULKULK ULK3 unc-51-like kinase 3 (C. elegans) 
501
ULK4OtherULKULK ULK4 unc-51-like kinase 4 (C. elegans) 
502
VACAMKLCAMKCAMK-Unique CAMKV CaM kinase-like vesicle-associated 
503
VRK1CK1VRK VRK1 vaccinia related kinase 1 
504
VRK2CK1VRK VRK2 vaccinia related kinase 2 
505
VRK3CK1VRK VRK3 vaccinia related kinase 3 
506
Wee1OtherWEE WEE1 WEE1 homolog (S. pombe) 
507
Wee1BOtherWEE FLJ40852 hypothetical protein FLJ40852 
508
Wnk1OtherWNK WNK1 WNK lysine deficient protein kinase 1 
509
Wnk2OtherWNK WNK2 WNK lysine deficient protein kinase 2 
510
Wnk3OtherWNK WNK3 WNK lysine deficient protein kinase 3 
511
Wnk4OtherWNK WNK4 WNK lysine deficient protein kinase 4 
512
YANK1AGCYANK STK32A serine/threonine kinase 32A 
513
YANK2AGCYANK STK32B serine/threonine kinase 32B 
514
YANK3AGCYANK STK32C serine/threonine kinase 32C 
515
YESTKSrcSrcA YES1 v-yes-1 Yamaguchi sarcoma viral oncogene homolog 1 
516
YSK1STESTE20YSK STK25 serine/threonine kinase 25 (STE20 homolog, yeast) 
517
ZAKTKLMLKZAK ZAK sterile alpha motif and leucine zipper containing kinase AZK 
518
ZAP70TKSyk ZAP70 zeta-chain (TCR) associated protein kinase 70kDa 


- Pseudokinases in human (Resource:Kinbase)

No.KinaseGroupFamilySubfamily
Pseudokinase 1
ARAFpsTKLRAFRAF 
Pseudokinase 2
AurAps1OtherAur 
Pseudokinase 3
AurAps2OtherAur 
Pseudokinase 4
AurBps1OtherAur 
Pseudokinase 5
BMPR1Aps1TKLSTKRType1 
Pseudokinase 6
BMPR1Aps2TKLSTKRType1 
Pseudokinase 7
BRAFpsTKLRAFRAF 
Pseudokinase 8
CDK4psCMGCCDKCDK4 
Pseudokinase 9
CDK5psCMGCCDKCDK5 
Pseudokinase 10
CDK7psCMGCCDKCDK7 
Pseudokinase 11
CDK8psCMGCCDKCDK8 
Pseudokinase 12
CGDpsRGCRGC 
Pseudokinase 13
CHK2ps1CAMKRAD53 
Pseudokinase 14
CHK2ps2CAMKRAD53 
Pseudokinase 15
CK1aps1CK1CK1CK1-A 
Pseudokinase 16
CK1aps2CK1CK1CK1-A 
Pseudokinase 17
CK1aps3CK1CK1CK1-A 
Pseudokinase 18
CK1g2psCK1CK1CK1-G 
Pseudokinase 19
CK2a1-rsCMGCCK2 
Pseudokinase 20
CLK2psCMGCCLK 
Pseudokinase 21
CLK3psCMGCCLK 
Pseudokinase 22
Erk3ps1CMGCMAPKERK3 
Pseudokinase 23
Erk3ps2CMGCMAPKERK3 
Pseudokinase 24
Erk3ps3CMGCMAPKERK3 
Pseudokinase 25
Erk3ps4CMGCMAPKERK3 
Pseudokinase 26
FERpsTKFer 
Pseudokinase 27
FLT1psTKVEGFR 
Pseudokinase 28
GPRK6psAGCGRKGRK 
Pseudokinase 29
HRIpsOtherPEKHRI 
Pseudokinase 30
KSGCpsRGCRGC 
Pseudokinase 31
LIMK2psTKLLISKLIMK 
Pseudokinase 32
MAP2K1psSTESTE7MEK1 
Pseudokinase 33
MAP2K2psSTESTE7MEK1 
Pseudokinase 34
MAPKAPKps1CAMKMAPKAPKMAPKAPK 
Pseudokinase 35
MARKps01CAMKCAMKLMARK 
Pseudokinase 36
MARKps02CAMKCAMKLMARK 
Pseudokinase 37
MARKps03CAMKCAMKLMARK 
Pseudokinase 38
MARKps04CAMKCAMKLMARK 
Pseudokinase 39
MARKps05CAMKCAMKLMARK 
Pseudokinase 40
MARKps07CAMKCAMKLMARK 
Pseudokinase 41
MARKps08CAMKCAMKLMARK 
Pseudokinase 42
MARKps09CAMKCAMKLMARK 
Pseudokinase 43
MARKps10CAMKCAMKLMARK 
Pseudokinase 44
MARKps11CAMKCAMKLMARK 
Pseudokinase 45
MARKps12CAMKCAMKLMARK 
Pseudokinase 46
MARKps13CAMKCAMKLMARK 
Pseudokinase 47
MARKps15CAMKCAMKLMARK 
Pseudokinase 48
MARKps16CAMKCAMKLMARK 
Pseudokinase 49
MARKps17CAMKCAMKLMARK 
Pseudokinase 50
MARKps18CAMKCAMKLMARK 
Pseudokinase 51
MARKps19CAMKCAMKLMARK 
Pseudokinase 52
MARKps20CAMKCAMKLMARK 
Pseudokinase 53
MARKps21CAMKCAMKLMARK 
Pseudokinase 54
MARKps22CAMKCAMKLMARK 
Pseudokinase 55
MARKps23CAMKCAMKLMARK 
Pseudokinase 56
MARKps24CAMKCAMKLMARK 
Pseudokinase 57
MARKps25CAMKCAMKLMARK 
Pseudokinase 58
MARKps26CAMKCAMKLMARK 
Pseudokinase 59
MARKps27CAMKCAMKLMARK 
Pseudokinase 60
MARKps28CAMKCAMKLMARK 
Pseudokinase 61
MARKps29CAMKCAMKLMARK 
Pseudokinase 62
MARKps30CAMKCAMKLMARK 
Pseudokinase 63
MNK1psCAMKMAPKAPKMNK 
Pseudokinase 64
MPSK1psOtherNAKMPSK 
Pseudokinase 65
MRCKpsAGCDMPKGEK 
Pseudokinase 66
MST3psSTESTE20YSK 
Pseudokinase 67
NEK2ps1OtherNEKNEK2 
Pseudokinase 68
NEK2ps2OtherNEKNEK2 
Pseudokinase 69
NEK2ps3OtherNEKNEK2 
Pseudokinase 70
NEK4psOtherNEKNEK4 
Pseudokinase 71
p70S6Kps1AGCRSKp70 
Pseudokinase 72
p70S6Kps2AGCRSKp70 
Pseudokinase 73
PAK2psSTESTE20PAKA 
Pseudokinase 74
PHKg1ps1CAMKPHK 
Pseudokinase 75
PHKg1ps2CAMKPHK 
Pseudokinase 76
PHKg1ps3CAMKPHK 
Pseudokinase 77
PKCipsAGCPKCIota 
Pseudokinase 78
PLK1ps1OtherPLKPLK1 
Pseudokinase 79
PLK1ps2OtherPLKPLK1 
Pseudokinase 80
PRKXpsAGCPKA 
Pseudokinase 81
PRP4psCMGCDYRKPRP4 
Pseudokinase 82
PSKH1psCAMKPSK 
Pseudokinase 83
RAF1psTKLRAFRAF 
Pseudokinase 84
RIOK3psAtypicalRIORIO3 
Pseudokinase 85
RYKpsTKRyk 
Pseudokinase 86
SAKpsOtherPLKSAK 
Pseudokinase 87
SCYL2psOtherSCY1 
Pseudokinase 88
SgK050psRGCRGC 
Pseudokinase 89
SgK384psOtherPLKPLK2 
Pseudokinase 90
SRPK2psCMGCSRPK 
Pseudokinase 91
STK33psCAMKCAMK-Unique 
Pseudokinase 92
STLK6ps1STESTE20STLK 
Pseudokinase 93
STLK6-rsSTESTE20STLK 
Pseudokinase 94
TLK1psOtherTLK 
Pseudokinase 95
TLK2ps1OtherTLK 
Pseudokinase 96
TLK2ps2OtherTLK 
Pseudokinase 97
TSSKps1CAMKTSSK 
Pseudokinase 98
TSSKps2CAMKTSSK 
Pseudokinase 99
TYRO3psTKAxl 
Pseudokinase 100
VRK3psCK1VRK 
Pseudokinase 101
Wee1BpsOtherWEE 
Pseudokinase 102
Wee1ps1OtherWEE 
Pseudokinase 103
Wee1ps2OtherWEE 
Pseudokinase 104
YESpsTKSrcSrcA